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A growth cost of begging in captive canary chicks. Proc. Natl. Acad. Sci. 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O;)3 d$5;,Ic*nWo¾g zK,㴰ӭR-Ph*@^ xo#-+Lӭc: Pq WxW@|+XxGi֑fj[(@TbWnQ8QȈ$t>  1\D́ҹ/|A>):~^YDPWQB![i*ňr^ w|"exKź7"Ye1V|n}qb]qi+HbY`(NsRQF3ARDXT69 ފG$u) Mɕ n"!Lvs@P;9]J=E`g4*T` (x|M?O|M;; %UK#'42e K?~x⟆5z~ZAmj#$J]*ɷM*":2b!So|G:žιym.+s}lO* Fue H=T1Gb4\ E`9'&B;(((((+ endstream endobj 88 0 obj << /Length 3078 /Subtype /XML /Type /Metadata >> stream 8 8 8 endstream endobj 89 0 obj [ 90 0 R 92 0 R 94 0 R 96 0 R 98 0 R 100 0 R 102 0 R 104 0 R 106 0 R 108 0 R 110 0 R 112 0 R 114 0 R 116 0 R 118 0 R 120 0 R 122 0 R 124 0 R ] endobj 90 0 obj << /Type /Annot /Subtype /Link /Rect [ 166.13 472.263 171.784 460.122 ] /BS << /S /S /Type /Border /W 0 >> /Border [ 0 0 0 ] /A 91 0 R /H /N >> endobj 91 0 obj << /S /GoTo /D (NPG_SREP_SREP40794.indd:Verhulst, S., Aviv, A., Benetos, A., Berenson, G. S. & Kark, J. D. Do leukocyte telomere length dynamics depend on baseline telomere length? An analysis that corrects for regression to the mean . Eur. J. 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Mean high sensitivity C-reactive protein \(HS-CRP; square-root  g/ml\) and interleukin-6 \(IL-6, log pg/ml\) in adult plasma, by experimental group. Error bars represent one standard error.:75) [ 76 0 R /XYZ 155 269 null ] (NPG_SREP_SREP40794.indd:Figure 5. Adult plasma interleukin-6 \(IL-6, log pg/ml\) against day 56 oxidative DNA damage \(pg 8-OHdG/g DNA\). The line represents a least-squares regression line and the shaded area its 95% confidence interval.:76) [ 85 0 R /XYZ 155 521 null ] (NPG_SREP_SREP40794.indd:Fisher, M. O., Nager, R. G. & Monaghan, P. Compensatory growth impairs adult cognitive performance. PLoS Biol. 4, e251 \(2006\).\r:186) [ 192 0 R /XYZ 179 173 null ] (NPG_SREP_SREP40794.indd:Fridsolfsson, A.-K. & Ellegren, H. A simple and universal method for molecular sexing of non-ratite birds. J. Avian Biol. 30, 116 21 \(1999\).\r:246) [ 194 0 R /XYZ 179 489 null ] (NPG_SREP_SREP40794.indd:Gavrilov, L. A. & Gavrilova, N. S. 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Sci. 276, 1679 83 \(2009\).\r:102) [ 192 0 R /XYZ 179 335 null ] (NPG_SREP_SREP40794.indd:Boonekamp, J. J., Mulder, G. A., Salomons, H. M., Dijkstra, C. & Verhulst, S. Nestling telomere shortening, but not telomere length, reflects developmental stress and predicts survival in wild birds. Proc. R. Soc. B-Bio:101) [ 192 0 R /XYZ 179 445 null ] (NPG_SREP_SREP40794.indd:Boonekamp, J. J., Simons, M. J. P., Hemerik, L. & Verhulst, S. Telomere length behaves as biomarker of somatic redundancy rather than biological age. Aging Cell 12, 330 2 \(2013\).\r:169) [ 192 0 R /XYZ 179 352 null ] (NPG_SREP_SREP40794.indd:Cawthon, R. M. Telomere measurement by quantitative PCR. Nucleic Acids Res. 30, 1 6 \(2002\).\r:237) [ 194 0 R /XYZ 179 549 null ] (NPG_SREP_SREP40794.indd:Danese, A., Pariante, C. M., Caspi, A., Taylor, A. & Poulton, R. Childhood maltreatment predicts adult inflammation in a life-course study. Proc. Natl. Acad. Sci. USA 104, 1319 24 \(2007\).\r:210) [ 192 0 R /XYZ 179 581 null ] (NPG_SREP_SREP40794.indd:Figure 1. Effects of experimental treatments on development. Error bars represent one standard error. \(a\) Mean daily weight by experimental group during and immediately after the manipulation. \(b\) Mean tarsus length by experimental:66) [ 54 0 R /XYZ 155 488 null ] (NPG_SREP_SREP40794.indd:Figure 2. Telomere dynamics. \(a\) Relationship between juvenile telomere length \(day 56\) and telomere length at day 5. The solid line represents x = y. If there were no overall attrition, observations would be expected to fall around:69) [ 69 0 R /XYZ 155 304 null ] (NPG_SREP_SREP40794.indd:Gavrilov, L. A. & Gavrilova, N. S. Early-Life Programming of Aging and Longevity: The Idea of High Initial Damage Load \(the HIDL Hypothesis\). Ann. N. Y. Acad. Sci. 1019, 496 501 \(2004\).\r:174) [ 192 0 R /XYZ 179 267 null ] (NPG_SREP_SREP40794.indd:Heidinger, B. J. et al. Telomere length in early life predicts lifespan. Proc. Natl. Acad. Sci. USA 109, 1743 8 \(2012\).\r:80) [ 192 0 R /XYZ 179 318 null ] (NPG_SREP_SREP40794.indd:Kuller, L. H., Tracy, R. P., Shaten, J. & Meilahn, E. N. Relation of C-Reactive Protein and Coronary Heart Disease in the MRFIT Nested Case-Control Study. American J. Epidemiol. 144, 537 547 \(1996\).\r:164) [ 192 0 R /XYZ 179 513 null ] (NPG_SREP_SREP40794.indd:Miller, G. E., Chen, E. & Parker, K. J. Psychological stress in childhood and susceptibility to the chronic diseases of aging: moving toward a model of behavioral and biological mechanisms. Psychol. Bull. 137, 959 97 \(2011:175) [ 192 0 R /XYZ 179 692 null ] (NPG_SREP_SREP40794.indd:Nettle, D. et al. An experimental demonstration that early-life competitive disadvantage accelerates telomere loss. Proc. R. Soc. B-Biological Sci. 282, 20141610 \(2015\).\r:120) [ 192 0 R /XYZ 179 411 null ] (NPG_SREP_SREP40794.indd:Nettle, D. et al. Brood size moderates associations between body size, telomere length and immune development in European starling nestlings. Ecol. Evol. online early access doi: 10.1002/ece3.2551 \(2016\).\r:121) [ 192 0 R /XYZ 179 156 null ] (NPG_SREP_SREP40794.indd:Nettle, D., Monaghan, P., Boner, W., Gillespie, R. & Bateson, M. Bottom of the Heap: Having Heavier Competitors Accelerates Early-Life Telomere Loss in the European Starling, Sturnus vulgaris. PLoS One 8, e83617 \(:98) [ 192 0 R /XYZ 179 428 null ] (NPG_SREP_SREP40794.indd:Nowicki, S., Searcy, W. A. & Peters, S. Brain development, song learning and mate choice in birds: a review and experimental test of the nutritional stress hypothesis . J. Comp. Physiol. A. Neuroethol. Sens. Neural. Behav. Physiol:184) [ 192 0 R /XYZ 179 190 null ] (NPG_SREP_SREP40794.indd:Shalev, I. et al. Stress and telomere biology: A lifespan perspective. Psychoneuroendocrinology 38, 1835 1842 \(2013\).\r:172) [ 192 0 R /XYZ 179 309 null ] (NPG_SREP_SREP40794.indd:Verhulst, S., Aviv, A., Benetos, A., Berenson, G. S. & Kark, J. D. Do leukocyte telomere length dynamics depend on baseline telomere length? An analysis that corrects for regression to the mean . Eur. J. Epidemiol. 2:202) [ 192 0 R /XYZ 179 139 null ] (NPG_SREP_SREP40794.indd:Bateson, M. Cumulative stress in research animals: Telomere attrition as a biomarker in a welfare context? BioEssays 38, 201 12 \(2016\).\r:185) [ 192 0 R /XYZ 179 301 null ] (NPG_SREP_SREP40794.indd:Belsky, D. W. et al. Quantification of biological aging in young adults. Proc. Natl. Acad. Sci. 112, E4104 4110 \(2015\).\r:230) [ 192 0 R /XYZ 179 54 null ] (NPG_SREP_SREP40794.indd:Bize, P., Criscuol, F., Metcalfe, N. B., Nasir, L. & Monaghan, P. Telomere dynamics rather than age predict life expectancy in the wild. Proc. R. Soc. B Biol. Sci. 276, 1679 83 \(2009\).\r:170) [ 192 0 R /XYZ 179 335 null ] (NPG_SREP_SREP40794.indd:Boonekamp, J. J., Mulder, G. A., Salomons, H. M., Dijkstra, C. & Verhulst, S. Nestling telomere shortening, but not telomere length, reflects developmental stress and predicts survival in wild birds. Proc. R. Soc. B-Bi:168) [ 192 0 R /XYZ 179 445 null ] (NPG_SREP_SREP40794.indd:Boonekamp, J. J., Simons, M. J. P., Hemerik, L. & Verhulst, S. Telomere length behaves as biomarker of somatic redundancy rather than biological age. Aging Cell 12, 330 2 \(2013\).\r:226) [ 192 0 R /XYZ 179 352 null ] (NPG_SREP_SREP40794.indd:Cawthon, R. M. Telomere measurement by quantitative PCR. Nucleic Acids Res. 30, 1 6 \(2002\).\r:239) [ 194 0 R /XYZ 179 549 null ] (NPG_SREP_SREP40794.indd:Figure 1. Effects of experimental treatments on development. Error bars represent one standard error. \(a\) Mean daily weight by experimental group during and immediately after the manipulation. \(b\) Mean tarsus length by experimenta:67) [ 54 0 R /XYZ 155 488 null ] (NPG_SREP_SREP40794.indd:Figure 2. Telomere dynamics. \(a\) Relationship between juvenile telomere length \(day 56\) and telomere length at day 5. The solid line represents x = y. If there were no overall attrition, observations would be expected to fall aroun:70) [ 69 0 R /XYZ 155 304 null ] (NPG_SREP_SREP40794.indd:Figure 3. Oxidative damage to DNA \(pg 8-OHdG/g DNA\) at day 56, by experimental group. Error bars represent one standard error.:197) [ 76 0 R /XYZ 155 520 null ] (NPG_SREP_SREP40794.indd:Figure 4. Inflammation markers in adulthood. Mean high sensitivity C-reactive protein \(HS-CRP; square-root  g/ml\) and interleukin-6 \(IL-6, log pg/ml\) in adult plasma, by experimental group. Error bars represent one standard error.:198) [ 76 0 R /XYZ 155 269 null ] (NPG_SREP_SREP40794.indd:Figure 5. Adult plasma interleukin-6 \(IL-6, log pg/ml\) against day 56 oxidative DNA damage \(pg 8-OHdG/g DNA\). The line represents a least-squares regression line and the shaded area its 95% confidence interval.:199) [ 85 0 R /XYZ 155 521 null ] (NPG_SREP_SREP40794.indd:Gavrilov, L. A. & Gavrilova, N. S. Early-Life Programming of Aging and Longevity: The Idea of High Initial Damage Load \(the HIDL Hypothesis\). Ann. N. Y. Acad. Sci. 1019, 496 501 \(2004\).\r:204) [ 192 0 R /XYZ 179 267 null ] (NPG_SREP_SREP40794.indd:Heidinger, B. J. et al. Telomere length in early life predicts lifespan. Proc. Natl. Acad. Sci. USA 109, 1743 8 \(2012\).\r:94) [ 192 0 R /XYZ 179 318 null ] (NPG_SREP_SREP40794.indd:Kuller, L. H., Tracy, R. P., Shaten, J. & Meilahn, E. N. Relation of C-Reactive Protein and Coronary Heart Disease in the MRFIT Nested Case-Control Study. American J. Epidemiol. 144, 537 547 \(1996\).\r:215) [ 192 0 R /XYZ 179 513 null ] (NPG_SREP_SREP40794.indd:Miller, G. E., Chen, E. & Parker, K. J. Psychological stress in childhood and susceptibility to the chronic diseases of aging: moving toward a model of behavioral and biological mechanisms. Psychol. Bull. 137, 959 97 \(2011:209) [ 192 0 R /XYZ 179 692 null ] (NPG_SREP_SREP40794.indd:Nettle, D. et al. An experimental demonstration that early-life competitive disadvantage accelerates telomere loss. Proc. R. Soc. B-Biological Sci. 282, 20141610 \(2015\).\r:183) [ 192 0 R /XYZ 179 411 null ] (NPG_SREP_SREP40794.indd:Nettle, D. et al. Brood size moderates associations between body size, telomere length and immune development in European starling nestlings. Ecol. Evol. 6, 8138 48 \(2016\).\r:201) [ 192 0 R /XYZ 179 156 null ] (NPG_SREP_SREP40794.indd:Nettle, D., Monaghan, P., Boner, W., Gillespie, R. & Bateson, M. Bottom of the Heap: Having Heavier Competitors Accelerates Early-Life Telomere Loss in the European Starling, Sturnus vulgaris. PLoS One 8, e83617 \(:114) [ 192 0 R /XYZ 179 428 null ] ] /Limits [ (NPG_SREP_SREP40794.indd:Lacey, R. E., Kumari, M. & McMunn, A. Parental separation in childhood and adult inflammation: the importance of material and psychosocial pathways. Psychoneuroendocrinology 38, 2476 84 \(2013\).\r:162) (NPG_SREP_SREP40794.indd:Nettle, D., Monaghan, P., Boner, W., Gillespie, R. & Bateson, M. Bottom of the Heap: Having Heavier Competitors Accelerates Early-Life Telomere Loss in the European Starling, Sturnus vulgaris. PLoS One 8, e83617 \(:114) ] >> endobj 245 0 obj << /Names [ (NPG_SREP_SREP40794.indd:Nowicki, S., Searcy, W. A. & Peters, S. Brain development, song learning and mate choice in birds: a review and experimental test of the nutritional stress hypothesis . J. Comp. Physiol. A. Neuroethol. Sens. Neural. Behav. 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J., Mulder, G. A., Salomons, H. M., Dijkstra, C. & Verhulst, S. Nestling telomere shortening, but not telomere length, reflects developmental stress and predicts survival in wild birds. Proc. R. Soc. B-B:176) [ 192 0 R /XYZ 179 445 null ] (NPG_SREP_SREP40794.indd:Figure 1. Effects of experimental treatments on development. Error bars represent one standard error. \(a\) Mean daily weight by experimental group during and immediately after the manipulation. \(b\) Mean tarsus length by experiment:68) [ 54 0 R /XYZ 155 488 null ] (NPG_SREP_SREP40794.indd:Figure 2. Telomere dynamics. \(a\) Relationship between juvenile telomere length \(day 56\) and telomere length at day 5. The solid line represents x = y. If there were no overall attrition, observations would be expected to fall arou:71) [ 69 0 R /XYZ 155 304 null ] (NPG_SREP_SREP40794.indd:Heidinger, B. J. et al. Telomere length in early life predicts lifespan. Proc. Natl. Acad. Sci. USA 109, 1743 8 \(2012\).\r:103) [ 192 0 R /XYZ 179 318 null ] (NPG_SREP_SREP40794.indd:Nettle, D. et al. An experimental demonstration that early-life competitive disadvantage accelerates telomere loss. Proc. R. Soc. B-Biological Sci. 282, 20141610 \(2015\).\r:221) [ 192 0 R /XYZ 179 411 null ] (NPG_SREP_SREP40794.indd:Nettle, D. et al. Brood size moderates associations between body size, telomere length and immune development in European starling nestlings. Ecol. Evol. 6, 8138 48 \(2016\).\r:222) [ 192 0 R /XYZ 179 156 null ] (NPG_SREP_SREP40794.indd:Nettle, D., Monaghan, P., Boner, W., Gillespie, R. & Bateson, M. Bottom of the Heap: Having Heavier Competitors Accelerates Early-Life Telomere Loss in the European Starling, Sturnus vulgaris. PLoS One 8, e83617 :119) [ 192 0 R /XYZ 179 428 null ] (NPG_SREP_SREP40794.indd:Bize, P., Criscuol, F., Metcalfe, N. B., Nasir, L. & Monaghan, P. Telomere dynamics rather than age predict life expectancy in the wild. Proc. R. Soc. B Biol. Sci. 276, 1679 83 \(2009\).\r:224) [ 192 0 R /XYZ 179 335 null ] (NPG_SREP_SREP40794.indd:Boonekamp, J. J., Mulder, G. A., Salomons, H. M., Dijkstra, C. & Verhulst, S. Nestling telomere shortening, but not telomere length, reflects developmental stress and predicts survival in wild birds. Proc. R. Soc. B-:223) [ 192 0 R /XYZ 179 445 null ] (NPG_SREP_SREP40794.indd:Figure 2. Telomere dynamics. \(a\) Relationship between juvenile telomere length \(day 56\) and telomere length at day 5. The solid line represents x = y. If there were no overall attrition, observations would be expected to fall aro:72) [ 69 0 R /XYZ 155 304 null ] (NPG_SREP_SREP40794.indd:Heidinger, B. J. et al. Telomere length in early life predicts lifespan. Proc. Natl. Acad. Sci. USA 109, 1743 8 \(2012\).\r:171) [ 192 0 R /XYZ 179 318 null ] (NPG_SREP_SREP40794.indd:Nettle, D. et al. An experimental demonstration that early-life competitive disadvantage accelerates telomere loss. Proc. R. Soc. B-Biological Sci. 282, 20141610 \(2015\).\r:242) [ 192 0 R /XYZ 179 411 null ] (NPG_SREP_SREP40794.indd:Nettle, D. et al. Brood size moderates associations between body size, telomere length and immune development in European starling nestlings. Ecol. Evol. 6, 8138 48 \(2016\).\r:243) [ 192 0 R /XYZ 179 156 null ] (NPG_SREP_SREP40794.indd:Nettle, D., Monaghan, P., Boner, W., Gillespie, R. & Bateson, M. Bottom of the Heap: Having Heavier Competitors Accelerates Early-Life Telomere Loss in the European Starling, Sturnus vulgaris. PLoS One 8, e83617:177) [ 192 0 R /XYZ 179 428 null ] (NPG_SREP_SREP40794.indd:Figure 1. Effects of experimental treatments on development. Error bars represent one standard error. \(a\) Mean daily weight by experimental group during and immediately after the manipulation. \(b\) Mean tarsus length by experim:187) [ 54 0 R /XYZ 155 488 null ] (NPG_SREP_SREP40794.indd:Figure 2. Telomere dynamics. \(a\) Relationship between juvenile telomere length \(day 56\) and telomere length at day 5. The solid line represents x = y. If there were no overall attrition, observations would be expected to fall ar:73) [ 69 0 R /XYZ 155 304 null ] (NPG_SREP_SREP40794.indd:Heidinger, B. J. et al. Telomere length in early life predicts lifespan. Proc. Natl. Acad. Sci. USA 109, 1743 8 \(2012\).\r:173) [ 192 0 R /XYZ 179 318 null ] (NPG_SREP_SREP40794.indd:Nettle, D., Monaghan, P., Boner, W., Gillespie, R. & Bateson, M. Bottom of the Heap: Having Heavier Competitors Accelerates Early-Life Telomere Loss in the European Starling, Sturnus vulgaris. PLoS One 8, e83617:179) [ 192 0 R /XYZ 179 428 null ] (NPG_SREP_SREP40794.indd:Figure 1. Effects of experimental treatments on development. Error bars represent one standard error. \(a\) Mean daily weight by experimental group during and immediately after the manipulation. \(b\) Mean tarsus length by experi:189) [ 54 0 R /XYZ 155 488 null ] (NPG_SREP_SREP40794.indd:Heidinger, B. J. et al. Telomere length in early life predicts lifespan. Proc. Natl. Acad. Sci. USA 109, 1743 8 \(2012\).\r:203) [ 192 0 R /XYZ 179 318 null ] (NPG_SREP_SREP40794.indd:Nettle, D., Monaghan, P., Boner, W., Gillespie, R. & Bateson, M. Bottom of the Heap: Having Heavier Competitors Accelerates Early-Life Telomere Loss in the European Starling, Sturnus vulgaris. PLoS One 8, e8361:220) [ 192 0 R /XYZ 179 428 null ] (NPG_SREP_SREP40794.indd:Figure 1. Effects of experimental treatments on development. Error bars represent one standard error. \(a\) Mean daily weight by experimental group during and immediately after the manipulation. \(b\) Mean tarsus length by exper:190) [ 54 0 R /XYZ 155 488 null ] (NPG_SREP_SREP40794.indd:Figure 2. Telomere dynamics. \(a\) Relationship between juvenile telomere length \(day 56\) and telomere length at day 5. The solid line represents x = y. If there were no overall attrition, observations would be expected to fall :188) [ 69 0 R /XYZ 155 304 null ] (NPG_SREP_SREP40794.indd:Heidinger, B. J. et al. Telomere length in early life predicts lifespan. Proc. Natl. Acad. Sci. USA 109, 1743 8 \(2012\).\r:217) [ 192 0 R /XYZ 179 318 null ] (NPG_SREP_SREP40794.indd:Nettle, D., Monaghan, P., Boner, W., Gillespie, R. & Bateson, M. Bottom of the Heap: Having Heavier Competitors Accelerates Early-Life Telomere Loss in the European Starling, Sturnus vulgaris. PLoS One 8, e836:236) [ 192 0 R /XYZ 179 428 null ] (NPG_SREP_SREP40794.indd:Figure 1. Effects of experimental treatments on development. Error bars represent one standard error. \(a\) Mean daily weight by experimental group during and immediately after the manipulation. \(b\) Mean tarsus length by expe:191) [ 54 0 R /XYZ 155 488 null ] (NPG_SREP_SREP40794.indd:Figure 2. Telomere dynamics. \(a\) Relationship between juvenile telomere length \(day 56\) and telomere length at day 5. The solid line represents x = y. If there were no overall attrition, observations would be expected to fall:192) [ 69 0 R /XYZ 155 304 null ] (NPG_SREP_SREP40794.indd:Heidinger, B. J. et al. Telomere length in early life predicts lifespan. Proc. Natl. Acad. Sci. USA 109, 1743 8 \(2012\).\r:225) [ 192 0 R /XYZ 179 318 null ] (NPG_SREP_SREP40794.indd:Nettle, D., Monaghan, P., Boner, W., Gillespie, R. & Bateson, M. Bottom of the Heap: Having Heavier Competitors Accelerates Early-Life Telomere Loss in the European Starling, Sturnus vulgaris. PLoS One 8, e83:241) [ 192 0 R /XYZ 179 428 null ] (NPG_SREP_SREP40794.indd:Figure 2. Telomere dynamics. \(a\) Relationship between juvenile telomere length \(day 56\) and telomere length at day 5. The solid line represents x = y. If there were no overall attrition, observations would be expected to fal:193) [ 69 0 R /XYZ 155 304 null ] (NPG_SREP_SREP40794.indd:Figure 2. Telomere dynamics. \(a\) Relationship between juvenile telomere length \(day 56\) and telomere length at day 5. The solid line represents x = y. If there were no overall attrition, observations would be expected to fa:194) [ 69 0 R /XYZ 155 304 null ] (NPG_SREP_SREP40794.indd:Figure 2. Telomere dynamics. \(a\) Relationship between juvenile telomere length \(day 56\) and telomere length at day 5. The solid line represents x = y. If there were no overall attrition, observations would be expected to f:195) [ 69 0 R /XYZ 155 304 null ] (NPG_SREP_SREP40794.indd:Figure 2. Telomere dynamics. \(a\) Relationship between juvenile telomere length \(day 56\) and telomere length at day 5. 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cellular ageing and affects adult inflammation: Experimental evidence from the European starling Daniel Nettle Clare Andrews Sophie Reichert Tom Bedford Claire Kolenda Craig Parker Carmen Martin-Ruiz Pat Monaghan Melissa Bateson Acrobat Distiller 11.0.9(Windows) © 2016 The Author(s) 10.1038/srep40794 2045-2322 Nature Publishing Group permissions@nature.com http://dx.doi.org/10.1038/srep40794 2017-01-07T16:58:11+05:30 2019-05-10T09:09:59+05:30 2019-05-10T09:09:59+05:30 doi:10.1038/srep40794 srep , (2016). doi:10.1038/srep40794 True uuid:2b2e2d10-4a0c-47ef-b3d1-93baf3f554c3 uuid:f87c38a4-2011-4fb2-876e-b46f238b41cf endstream endobj xref 0 268 0000000000 65535 f 0000136132 00000 n 0000136663 00000 n 0000147999 00000 n 0000148649 00000 n 0000148789 00000 n 0000148974 00000 n 0000149133 00000 n 0000149516 00000 n 0000149675 00000 n 0000150064 00000 n 0000150225 00000 n 0000150747 00000 n 0000150907 00000 n 0000151512 00000 n 0000151673 00000 n 0000152276 00000 n 0000152437 00000 n 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